Torosaurus Is Not Triceratops: Ontogeny in Chasmosaurine Ceratopsids as a Case Study in Dinosaur Taxonomy. 2012. N.R. Longrich and D. J. Field. PLoS ONE 7(2): e32623.
From PLoS One e32623[edit]: It has been proposed that the genera
Triceratops and
Torosaurus are in synonymous, with specimens identified as
Torosaurus representing the adult form of
Triceratops. The hypothesis of synonymy makes three testable predictions: 1) the species in question should have similar geographic and stratigraphic distributions, 2) specimens assigned to
Torosaurus should be more mature than those assigned to
Triceratops, and 3) intermediates should exist that combine features of Triceratops and
Torosaurus. The first condition appears to be met, but it remains unclear whether the other predictions are borne out by the fossil evidence.
We assessed the relative maturity of
Torosaurus and
Triceratops specimens by coding skulls for characters that vary with maturity, and then using a clustering analysis to arrange them into a growth series. We found that a well-defined sequence of changes exists in horned dinosaurs: development of cranial ornament occurs in juveniles, followed by fusion of the skull roof in subadults, and finally, the epoccipitals, epijugals, and rostral fuse to the skull in adults. Using this scheme, we identified mature and immature individuals of both
Torosaurus and
Triceratops. Furthermore, we describe the ventral depressions on the frill of Triceratops, and show that they differ in shape and position from the parietal fenestrae of
Torosaurus. Thus, we conclude that these structures are not intermediates between the solid frill of Triceratops and the fenestrated frill of
Torosaurus.