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began his career with T.H. Morgan studying mutations in fruit flies. He was the first to increase the mutation rate using heat, later using 50 kilovolt X-rays to induce an even greater incidence of mutations. Muller was awarded the Nobel Prize for Physiology or Medicine in 1946Muller long warned about needless exposures to radiation and their associated risks of cancer and heritable genetic effects. By the late 1940s, the nuclear weapons testing program had begun and Muller was a vocal critic of the Atomic Energy Commission's views on the hazards of worldwide fallout.
